Orthalicidae (orthalicid land snails) are a family of tropical air-breathing land snails, terrestrial pulmonate gastropod mollusks are classified in the superfamily Orthalicoidea of the order Stylommatophora.MolluscaBase eds. (2021). MolluscaBase. Orthalicidae Martens, 1860. Accessed through: World Register of Marine Species at: http://marinespecies.org/aphia.php?p=taxdetails&id=870044 on 2021-02-19
They are medium-sized to large snails, from about 3 cm (about 1.2 inches) to 9 cm (about 3.5 inches) in shell length
Taxonomy In former times, this family was often known as the Bulimulidae, but this term may also denote what today is the subfamily Bulimulinae. The subfamily Bulimulinae replaces the former family Bulimulidae Crosse & P. Fischer, 1873.
2005 taxonomy The Orthalicidae belong to the Orthalicoidea, a superfamily in the order Sigmurethra. Like other stylommatophorans, the Sigmurethra belong to the suborder Helicina. Among the three subfamilies of Orthalicidae, the Bulimulinae are the most diverse, consisting of three tribes:
subfamily Amphibuliminae P. Fischer, 1873 - synonym Peltellinae Gray, 1855 (check decision of International Commission on Zoological Nomenclature for Peltellinae) - 17 speciesBreure A., Groenenberg D. & Schilthuizen M. (2010). "Gondwana revisited: New insights in the phylogenetic relationships within the Orthalicoidea (Mollusca, Stylommatophora)". poster. PDF . subfamily Bulimulinae Tryon, 1867 - 1243 species tribe Bulimulini Tryon, 1867 - synonyms: Bulimidae Guilding, 1828 (inv.); Berendtiinae P. Fischer & Crosse, 1872; Bothiembryontidae Iredale, 1937 tribe Odontostomini Pilsbry & Vanatta, 1898 - 125 species tribe Simpulopsini Schileyko, 1999 - synonym: Tomogeridae Jousseaume, 1877 subfamily Orthalicinae Albers, 1850 - synonym: Liguidae Pilsbry, 1891 - 83 species
2010 taxonomy Breure et al. (2010) elevated Bulimulinae to Bulimulidae, Odontostomini to Odontostomidae, Amphibuliminae to Amphibulimidae.Breure A. S. H., Groenenberg D. S. J. & Schilthuizen M. (2010). "New insights in the phylogenetic relations within the Orthalicoidea (Gastropoda, Stylommatophora) based on 28S sequence data". Basteria 74(1-3): 25-31.
2012 taxonomy Breure & Romero (2012) confirmed previous results from 2010, additionally they elevated Simpulopsini to Simpulopsidae.Breure A. S. H. & Romero P. (2012). "Support and surprises: molecular phylogeny of the land snail superfamily Orthalicoidea using a three-locus gene analysis with a divergence time analysis and ancestral area reconstruction (Gastropoda: Stylommatophora)". Archiv für Molluskenkunde: International Journal of Malacology 141(1): 1-20. .
Genera Genera within the family Orthalicidae include: Aposcutalus Dutra & Leme, 1985 Clathrorthalicus Strebel, 1909 Corona Albers, 1850Breure A. S. H. & Mogollón Avila V. (2010). "Well-known and little-known: miscellaneous notes on Peruvian Orthalicidae (Gastropoda, Stylommatophora)". Zoologische Mededelingen 84. HTM . † Cortana Salvador & Simone, 2013 Hemibulimus Martens, 1885 Kara Strebel, 1910Breure A. S. H. (2011) "Annotated type catalogue of the Orthalicoidea (Mollusca, Gastropoda) in the Royal Belgian Institute of Sciences, Brussels, with descriptions of two new species". ZooKeys 101: 1–50. . Liguus Montfort, 1810 Orthalicus Beck, 1837 - type genus of the family Orthalicidae Paeniscutalus Wurtz, 1947 Porphyrobaphe Shuttleworth, 1856 Quechua Strebel, 1910 Scholvienia Strebel, 1910 Sultana Shuttleworth, 1856
References
Further reading Neubert E. & Janssen R. (2004). "Die Typen und Typoide des Natur-Museums Senckenberg, 84: Mollusca: Gastropoda: Pulmonata: Orthalicoidea: Bulimulidae (2), Orthalicidae, Placostylidae". Archiv für Molluskenkunde 133: 193–297. abstract. Ramirez J. & Ramírez R. (2010). "Análisis de la estructura secundaria del LSU rRNA mitocondrial de caracoles terrestres peruanos (Orthalicidae: Gastropoda) / Analysis of the secondary structure of mitochondrial LSU rRNA of Peruvian land snails (Orthalicidae: Gastropoda)." Revista Peruana de Biología 17(1): 53–57. PDF. Ramirez J., Ramírez R., Romero P., Chumbe A. & Ramírez P. (2009). "Posición evolutiva de caracoles terrestres peruanos (Orthalicidae) entre los Stylommatophora (Mollusca: Gastropoda) / Evolutionary position of Peruvian land snails (Orthalicidae) among Stylommatophora (Mollusca: Gastropoda)". Revista Peruana de Biología 16(1): 51–56. PDF.
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